As Adm.Lee stated... The USAF/USN officers would retain their rank because the only way to reduce an officer's rank is court-martials if i remember right. But they would be considered 'staff' officers... sort of like a IG, JAG, Medical Corps or Chaplin Corps until they got the training to be line officers (ie infantry, armor, arty, ect.).
The Army has what i think is called a 'Warrior Course' for prior service USAF and USN enlisted personnel who are transfering over to the US Army. I think it's a four week training course.
I have a feeling that these Officers would go through a crash course on infantry tactics, such as given to support services enlisted personnel to fill out combat arms as seen in the reorganization of the 5ID article I found online. In that article it said that enlisted support personnel where given crash course on infantry operations, and officers where trained to take over their jobs in the support services (i guess you could say that USAF Lt. Colonel is now serving food).
But in my campaign setting, we had done something a little different. The USAF created "guntruck squadrons" for pilots and flight crew who no longer had aircraft to fly or maintain. these 'guntruck sqadrons' would be assigned to rear echelon convoy escort and security enforcement duties.
While the USN created Naval Infantry Battalions that trained both officers and enlisted to fight as a cohesive unit. The navy had thse battalions being trained by USN SEALs and USMC... I recently discovered that the USN is working at creating Naval Infantry Battalions in real life using those men who 'washed out' of the SEALs program... I dont know if it is true, if not it really should be.
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